The literature of paleobiology is brimming with qualifiers and cautions about using species in the fossil record, or equating such species with those recognized among living organisms. Species and Speciation in the Fossil Record digs through this literature and surveys the recent research on species in paleobiology. In these pages, experts in the field examine what they think species are - in their particular taxon of specialty or more generally in the fossil record. They also reflect on what the answers mean for thinking about species in macroevolution. The first step in this approach is an overview of the Modern Synthesis, and paleobiology’s development of quantitative ways of documenting and analyzing variation with fossil assemblages. Following that, this volume’s central chapters explore the challenges of recognizing and defining species from fossil specimens, and show how with careful interpretation and a clear species concept, fossil species may be sufficiently robust for meaningful paleobiological analyses. Tempo and mode of speciation over time are also explored, exhibiting how the concept of species, if more refined, can reveal enormous amounts about the interplay between species origins and extinction and local and global climate change.

A collection of case studies that seeks to reexamine the understanding of the speciation patterns that appear in the fossil record through an analysis of the patterns and their presumed processes. In each case, the rigorous techniques of morphological analysis, quantitative genetic analysis, phylogenetic analysis, and sedimentary completeness have been employed.

The Principles of Biology sequence (BI 211, 212 and 213) introduces biology as a scientific discipline for students planning to major in biology and other science disciplines. Laboratories and classroom activities introduce techniques used to study biological processes and provide opportunities for students to develop their ability to conduct research.

The Paleobiological Revolution chronicles the incredible ascendance of the once-maligned science of paleontology to the vanguard of a field. With the establishment of the modern synthesis in the 1940s and the pioneering work of George Gaylord Simpson, Ernst Mayr, and Theodosius Dobzhansky, as well as the subsequent efforts of Stephen Jay Gould, David Raup, and James Valentine, paleontology became embedded in biology and emerged as paleobiology, a first-rate discipline central to evolutionary studies. Pairing contributions from some of the leading actors of the transformation with overviews from historians and philosophers of science, the essays here capture the excitement of the seismic changes in the discipline. In so doing, David Sepkoski and Michael Ruse harness the energy of the past to call for further study of the conceptual development of modern paleobiology.

Evolution is the central theme of all biology. Researcarcch in the many branches of evolutionary study continues to flourish. This book, based on a symposium of the Linnean Society, discusses the diversity in currentevolutionary research. It approaches the subject ambitiously and from several angles, bringing ttogether eminent authors from a variety of disciplines paleontologists traditionally with a macroevolutionary bias, neontologists concentrating on microevolutionary processes, and those studying the very essence ofsses and those studying the very essence of evolution the process of speciation in living organisms. Evolutionary Patterns and Processes will appeal to a broad spectrum of professional biologistsworking in such fields as paleontology, population biology, and evolutionary genetics. Biologists will enjoy chapters by Stephen J. Gould, discovering in the much earlier work of Hugo de Vries parallels with his ideas on punctuational evolution; Guy Bush,considering why there are so many small animals; Peter Sheldon, examining detailed fossil trilobite sequences for evidence of microevolutionary processes and considering models of speciation; as well as others dealing with cytological, ecological, and behavioral processes leading to the evolution of new species. None

The world’s most revered and eloquent interpreter of evolutionary ideas offers here a work of explanatory force unprecedented in our time—a landmark publication, both for its historical sweep and for its scientific vision. With characteristic attention to detail, Stephen Jay Gould first describes the content and discusses the history and origins of the three core commitments of classical Darwinism: that natural selection works on organisms, not genes or species; that it is almost exclusively the mechanism of adaptive evolutionary change; and that these changes are incremental, not drastic. Next, he examines the three critiques that currently challenge this classic Darwinian edifice: that selection operates on multiple levels, from the gene to the group; that evolution proceeds by a variety of mechanisms, not just natural selection; and that causes operating at broader scales, including catastrophes, have figured prominently in the course of evolution. Then, in a stunning tour de force that will likely stimulate discussion and debate for decades, Gould proposes his own system for integrating these classical commitments and contemporary critiques into a new structure of evolutionary thought. In 2001 the Library of Congress named Stephen Jay Gould one of America’s eighty-three Living Legends—people who embody the “quintessentially American ideal of individual creativity, conviction, dedication, and exuberance.” Each of these qualities finds full expression in this peerless work, the likes of which the scientific world has not seen—and may not see again—for well over a century.

A world of categones devmd of spirit waits for life to return. Saul Bellow, Humboldt's Gift The stock-in-trade of communicating hypotheses about the historical path of evolution is a graphical representation called a phylogenetic tree. In most such graphics, pairs of branches diverge from other branches, successively marching across abstract time toward the present. To each branch is tied a tag with a name, a binominal symbol that functions as does the name given to an individual human being. On phylogenetic trees the names symbolize species. What exactly do these names signify? What kind of information is communicated when we claim to have knowledge of the following types? "Tetonius mathewzi was ancestral to Pseudotetonius ambiguus. " "The sample of fossils attributed to Homo habzlis is too variable to contain only one species. " "Interbreeding populations of savanna baboons all belong to Papio anubis. " "Hylobates lar and H. pileatus interbreed in zones of geographic overlap. " While there is nearly universal agreement that the notion of the speczes is fundamental to our understanding of how evolution works, there is a very wide range of opinion on the conceptual content and meaning of such particular statements regarding species. This is because, oddly enough, evolutionary biolo gists are quite far from agreement on what a species is, how it attains this status, and what role it plays in evolution over the long term.

This 3-volume handbook brings together contributions by the world ́s leading specialists that reflect the broad spectrum of modern palaeoanthropology, thus presenting an indispensable resource for professionals and students alike. Vol. 1 reviews principles, methods, and approaches, recounting recent advances and state-of-the-art knowledge in phylogenetic analysis, palaeoecology and evolutionary theory and philosophy. Vol. 2 examines primate origins, evolution, behaviour, and adaptive variety, emphasizing integration of fossil data with contemporary knowledge of the behaviour and ecology of living primates in natural environments. Vol. 3 deals with fossil and molecular evidence for the evolution of Homo sapiens and its fossil relatives.

With all the recent advances in molecular and evolutionary biology, one could almost wonder why we need the fossil record. Molecular sequence data can resolve taxonomic relationships, experiments with fruit flies demonstrate evolution and development in real time, and field studies of Galapagos finches have provided the strongest evidence for natural selection ever measured in the wild. What, then, can fossils teach us that living organisms cannot? Evolutionary Patterns demonstrates the rich variety of clues to evolution that can be gleaned from the fossil record. Chief among these are the major trends and anomalies in species development revealed only by "deep time," such as periodic mass extinctions and species that remain unchanged in form for millions of years. Contributors explore modes of development, the tempo of speciation and extinction, and macroevolutionary patterns and trends. The result is an important contribution to paleobiology and evolutionary biology, and a spirited defense of the fossil record as a crucial tool for understanding evolution and development. The contributors are Ann F. Budd, Efstathia Bura, Leo W. Buss, Mike Foote, Jörn Geister, Stephen Jay Gould, Eckart Hâkansson, Jean-Georges Harmelin, Lee-Ann C. Hayek, Jeremy B. C. Jackson, Kenneth G. Johnson, Nancy Knowlton, Scott Lidgard, Frank K. McKinney, Daniel W. McShea, Ross H. Nehm, Beth Okamura, John M. Pandolfi, Paul D. Taylor, and Erik Thomsen.